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===== 2.4.3.2.1 Observed biome shifts predominantly driven by climate change ===== <div id="h4-15-siblings" class="h4-siblings"></div> AR5 and a meta-analysis found that vegetation at the biome level shifted poleward latitudinally and upward altitudinally due to anthropogenic climate change in at least 19 sites in boreal, temperate and tropical ecosystems from 1700 to 2007 ( [[#Gonzalez--2010|Gonzalez et al., 2010]] ; [[#Settele--2014|Settele et al., 2014]] ). In these areas, temperature increased to 0.4Β°Cβ1.6Β°C above the pre-industrial period ( [[#Gonzalez--2010|Gonzalez et al., 2010]] ; [[#Settele--2014|Settele et al., 2014]] ). Field research since the AR5 detected additional poleward and upslope biome shifts over periods of 24β210 years at numerous sites (described below), but were not directly attributed to anthropogenic climate change as the studies were not designed or conducted properly for full attribution assessment. Many of the recently detected shifts are nevertheless consistent with climate change-induced temperature increases and observed in areas without agriculture, livestock grazing, timber harvesting and other anthropogenic land uses. For example, in the Andes Mountains in Ecuador, a biome shift was detected by comparing a survey by Alexander von Humboldt in 1802 to a re-survey in 2012, making this the longest time span in the world for this type of data ( [[#Morueta-Holme--2015|Morueta-Holme et al., 2015]] ; [[#Moret--2019|Moret et al., 2019]] ). Over 210 years, temperature increased by 1.7Β°C ( [[#Morueta-Holme--2015|Morueta-Holme et al., 2015]] ) and the upper edge of alpine grassland shifted 100β450 m upslope ( [[#Moret--2019|Moret et al., 2019]] ). Other biome shifts consistent with climate change and not substantially affected by local land use include: northward shifts in Canada of deciduous forest into boreal conifer forest, 5 km from 1970β2012 ( [[#Sittaro--2017|Sittaro et al., 2017]] ) and 20 km from 1970β2014 ( [[#Boisvert-Marsh--2019|Boisvert-Marsh et al., 2019]] ) and of temperate conifer into boreal conifer forest, 21 km from 1970β2015 ( [[#Boisvert-Marsh--2021|Boisvert-Marsh and de Blois, 2021]] ). Research detected upslope shifts of boreal and sub-alpine conifer forest into alpine grassland at 143 sites on four continents (41 m from 1901β2018) ( [[#Lu--2021|Lu et al., 2021]] ) and at individual sites in Canada (54 m from 1900β2010) ( [[#Davis--2020|Davis et al., 2020]] ); China (300 m from 1910β2000) ( [[#Liang--2016|Liang et al., 2016]] ) (33 m from 1985β2014) ( [[#Du--2018|Du et al., 2018]] ); Nepal (50 m from 1860β2000) ( [[#Sigdel--2018|Sigdel et al., 2018]] ); Russia (150 m from 1954β2006) ( [[#Gatti--2019|Gatti et al., 2019]] ); and the USA (19 m from 1950β2016) ( [[#Smithers--2018|Smithers et al., 2018]] ) (38 m from 1953β2015) ( [[#Terskaia--2020|Terskaia et al., 2020]] ). Other upslope cases include shifts of temperate conifer forest in Canada ( [[#Jackson--2016|Jackson et al., 2016]] ) and the USA ( [[#Lubetkin--2017|Lubetkin et al., 2017]] ), temperate deciduous forest in Switzerland ( [[#Rigling--2013|Rigling et al., 2013]] ) and temperate shrubland in the USA ( [[#Donato--2016|Donato et al., 2016]] ). In summary, anthropogenic climate change caused latitudinal and elevational biome shifts in at least 19 sites in boreal, temperate and tropical ecosystems between 1700 and 2007, where temperature increased to 0.4Β°Cβ1.6Β°C above the pre-industrial period ( ''robust evidence'' , ''high agreement'' ). Additional cases of 5β20 km northward and 20β300 m upslope biome shifts between 1860 and 2016, under a mean global temperature increase of approximately 0.9Β°C above the pre-industrial period, are consistent with climate change ( ''medium evidence'' , ''high agreement'' ). <div id="2.4.3.2.2" class="h4-container"></div> <span id="observed-biome-shifts-from-combined-land-use-change-and-climate-change"></span>
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